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Bonebed Reference shelf for prehistoric life
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Dinosauria 17

Dinosaurs proper, including the birds that survived them.

  • Theropoda 9 Bipedal, mostly carnivorous, hollow-boned.
  • Sauropodomorpha 3 Long-necked, small-headed, graviportal.
  • Ornithischia 5 Bird-hipped plant eaters: plated, armoured, horned, beaked.

Reptiles (non-dinosaur) 4

Animals routinely filed next to dinosaurs and just as routinely mislabelled as one.

  • Pterosauria 1 Flying archosaurs with a membrane wing on one finger.
  • Mosasauridae 1 Open-ocean squamates with a flipper-paddled tail.
  • Plesiosauria 1 Four-paddle swimmers with a rigid trunk.
  • Ichthyosauria 1 Dolphin-shaped, live-bearing, ringed vertebrae.

Synapsida & mammals 2

The line that led to us. The sail-backs are in here, not with the dinosaurs.

  • Sphenacodontidae 1 Permian predators with a neural-spine sail.
  • Cetacea 0 Fully aquatic mammals with hind limbs reduced to vestiges.
  • Proboscidea 1 Elephants and their extinct kin; tusks are modified incisors.
  • Carnivora 0 Shearing carnassial teeth, four or five toes, no hooves.

Stem vertebrates 1

Jawed and jawless fish that carry the earliest bone in the record.

  • Placodermi 1 Arthrodire and antiarch head shields with a neck joint.

Invertebrates 3

Everything with an exoskeleton or no skeleton at all. Most of the fossil record by count.

  • Radiodonta 1 Stem arthropods with a ring of toothed grasping appendages.
  • Lobopodia 0 Soft-bodied panarthropods walking on stubby unjointed legs.
  • Trilobita 1 Calcite eyes, a three-lobed axis, and moult halves everywhere.
  • Insecta 1 Six legs, two wing pairs, tracheal respiration.
中文

Cretaceous 145 to 66.0 Ma

Triceratops

三角龙属

The most abundant large herbivore in the Hell Creek Formation, and the dinosaur whose skull is easiest to identify from a single fragment. Skulls reach 2 metres, a quarter of which is frill.

Published Reading time 1 min read

Triceratops — bone plate
Plate drawn from a per-clade shape library; scaled to published body length.

A skull built from fused plates

The frill is the parietal and squamosal bones extended backwards and locked together. In Triceratops it is solid, with no openings, which is unusual among ceratopsids and makes the skull heavy. The beak is a single bone at the front of the lower jaw, and the teeth sit in a shearing battery that replaced worn teeth continuously.

What it ate, and how we know

The lower jaw describes a near-vertical shear, and microwear on the teeth is dominated by fine parallel scratches rather than pits. That combination fits shearing tough, fibrous plants close to the ground -- ferns, cycads and early flowering shrubs -- rather than grazing abrasive grass, which had not yet spread.

  • Beak: cropping, not chewing.
  • Dental battery: up to 36 tooth positions per side, stacked.
  • Microwear: fine parallel striations, few pits.

Horn growth over a lifetime

Juveniles have short, straight brow horns that point upward, and the frill is proportionally small. Study of horn cross-sections shows they grew continuously and were probably covered in keratin, so their true length in life was longer than the bone core.

Evidence of what attacked it

Triceratops specimens carry a notable range of healed and unhealed injuries: punctures matching tyrannosaur teeth on frills and on the squamosal, bite marks on a horn, and one specimen with a partial tyrannosaur tooth embedded in a vertebra. There is also a documented infection around a broken horn, which suggests the horns were used in contact.

Injury type Element Reading
Puncture frill / squamosal tyrannosaur-sized tooth spacing
Healed fracture brow horn contact between individuals
Embedded tooth vertebra survived at least one attack

The three injury patterns most often cited from Hell Creek material.

Sources

Reference works and further reading for the material in this entry.

Where the argument is

What the frill is for. Sexual display, species recognition and thermoregulation have all been proposed. A 2022 review of frill blood-vessel traces found no convincing evidence of a heat-exchange role, which leaves display and species recognition as the surviving explanations rather than the proven ones.

Frequently misread

  • ClaimThe three horns point forward like a bull's.

    What the record showsThe brow horns sit above the eyes and curve forward and slightly outward; the third horn is a short, stout nasal horn. The frill, not the horns, is what dominates the skull from above.

  • ClaimIt is the ancestor of the rhinoceros.

    What the record showsTriceratops is an ornithischian dinosaur that died out 66 million years ago. Rhinoceroses are perissodactyl mammals with no dinosaur ancestry; the resemblance is convergence on the same low-browsing body plan.

Same period, same cabinet Cretaceous

  1. Tyrannosaurus 5.4 t - 8.8 t
  2. Ankylosaurus 4.5 t - 8.0 t
  3. Protoceratops 80 kg - 180 kg
  4. Therizinosaurus 3.0 t - 5.0 t